Lithophaga lithophaga (Linnaeus, 1758)
Southern coasts of the Bay of Biscay to Angola, Mediterranean. Shallow water to 25m deep.
 
Lives in colonies, nested inside walls, cliffs and even in human built structures, whenever the rock is of limestone. A famous example was given by the geologist Charles Lyell, who observed, on the three standing columns of the “Temple of Serapis” in Pozzuoli (pictured at left, with link), a band of drilled marble between 3m and 6m hight. The cause was a colony of L. lithophaga located just below the sea-level, at a time when the temple was partially submerged. The base of the columns remained intact, hidden and protected by volcanic deposits, nowadays removed. « Above this is a zone, about nine feet in height, where the marble has been pierced by a species of marine perforating bivalve, Lithodomus. » – C. Lyell: Principles of geology vol. I, London 1837.

Original taxon: Mytilus lithophagus. Synonyms: Lithodomus dactylus and inflatus, Lithophaga mytuloides.
2m deep, in limestone bank, Marathi beach, Souda bay, Chania, N. Crete. 37mm.
Baia di Giglio Campese, Isola del Giglio, Toscana, W. Italy. Original picture provided by V. Moser for iNaturalist.
– (CC BY) –
Unlike the animals of the genus Pholas, which perforate mechanically, the species of the genus Lithophaga are chemical drillers. Beach drift, Cap d’Agde, S. France. 47mm.
In rocky wall, northwestern coast of Krk island, Primorje-Gorski Kotar Comitat, N. Croatia. Original picture provided by S. Pav for iNaturalist – (CC BY-NC).
1m deep, Águilas, Murcia, S. Spain. 59-78mm.
Mytilus lithophagus in F. Berge: Conchylienbuch, Stuttgart 1855, via BHL.
Lithophaga lithoglypha in W. Dunker: “Die Gattung Lithophaga”, Systematisches Conchylien-Cabinet Bd.8:Abt.3a, Nürnberg 1882, via BHL.
 
Lithophaga lithoglypha Meuschen. – Testa valde inflata, solida, subcylindrica, utrinque rotundata, postice latior, striis incrementi concentricis subtilibus partem posticam versus fortioribus instructa ; basis paene rectilinea ; dorsum medium paululum fornicatum et carinatum ; umbones ab extremitate antica remotiusculi ; epidermis castanea, parte antica et dorso exceptis striis directis insignita. – Mare mediterraneum. Multis locis corallia et rupes perforans invenitur. »
Beachstormed specimen, Cádiz area, SW. Spain.
Original picture provided by B. J. Muñoz-Sánchez (ES).
– (CC BY-NC-SA) –
Other beachstormed specimen from the same coast. 39,6mm.
Original picture provided by B. J. Muñoz-Sánchez (ES).
– (CC BY-NC-SA) –
« Un pezzo di scoglio, ove sono annicchiati due Mitili Litofagi, ossia Dattili di mare […] vi è in oltre una sezione […] di una di siffatte nicchie. » – G. S. Poli: Testacea utriusque Siciliae… vol. II, Parma 1795 p. xxxvii and in the Atlas pl. XXXII.

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